Southern
Research Station

Education

M.S. in Entomology, 2000

University of Georgia

B.S. in Wildlife Science, 1997

Auburn University

Professional Experience

Entomologist, U.S. Forest Service

2001—Current

Featured Publications and Products

• Hanula, James L.; Horn, Scott; O'Brien, Joseph J. 2015. Have changing forests conditions contributed to pollinator decline in the southeastern United States?.
• Hanula, James; Horn, Scott. 2011. Removing an exotic shrub from riparian forests increases butterfly abundance and diversity.
• Horn, Scott; Hanula, James. 2011. Influence of trap color on collection of the recently introduced Bean Plataspid, Megacopta cribraria (Hemiptera: Plataspidae).
• Hanula, James; Ulyshen, Michael; Horn`, Scott. 2011. Effect of trap type, trap position, time of year, and beetle density on captures of the Redbay Ambrosia Beetle (Coleoptera: Curculionidae: Scolytinae).
• Ulyshen, Michael D.; Horn, Scott; Hanula, James L. 2010. Response of beetles (Coleoptera) at three heights to the experimental removal of an invasive shrub, Chinese privet (Lingustrum sinense), from floodplain forests.
• Ulyshen, Michael D.; Horn, Scott; Barnes, Brittany; Gandhi, Kamal J. K. 2010. Impacts of prescribed fire on saproxylic beetles in loblolly pine logs.
• Hudson, Jacob R.; Hanula, James; Horn, Scott. 2014. Removing Chinese privet from riparian forests still benefits pollinators five years later.
• Hudson, Jacob R.; Hanula, James L.; Horn, Scott. 2014. Impacts of removing Chinese privet from riparian forests on plant communities and tree growth five years later.
• Zhang, Yanzhuo; Hanula, James L.; Horn, Scott. 2012. The biology and preliminary host range of Megacopta cribraria (Heteroptera: Plataspidae) and its impact on kudzu growth.
• Hanula, James L.; Horn, Scott. 2011. Removing an invasive shrub (Chinese privet) increases native bee diversity and abundance in riparian forests of the southeastern United States.

Publications

• Ulyshen, Michael ; Horn, Scott . 2023. Declines of bees and butterflies over 15 years in a forested landscape.
• Ulyshen, Michael D.; Horn, Scott ; Hanula, James L. 2022. Decadal patterns of forest and pollinator recovery following the eradication of an invasive shrub.
• Vogt, James T.; MacGown, Joe A.; Lewis, James ; Horn, Scott ; Ulyshen, Michael D. 2022. Diversity and seasonal occurrence of native and nonnative ants (Hymenoptera: Formicidae) in long-term experimental Chinese Privet (Lamiales: Oleaceae) plots in Georgia, USA.
• Ulyshen, Michael ; Elliott, Katherine ; Scott, Joel ; Horn, Scott ; Clinton, Patsy ; Liu, Ning ; Miniat, Chelcy F.; Caldwell, Peter ; Oishi, Chris ; Knoepp, Jennifer ; Bolstad, Paul . 2022. Effects of Rhododendron removal and prescribed fire on bees and plants in the southern Appalachians.
• Poole, Emilee ; Ulyshen, Michael ; Horn, Scott ; Anderson, Patrick ; Bates, Chip ; Barnes, Chris . 2021. Progression of sugarberry ( Celtis laevigata ) dieback and mortality in the southeastern United States.
• Ulyshen, Michael D.; Horn, Scott ; Brownie, Cavell ; Strickland, Michael S.; Wurzburger, Nina ; Zanne, Amy . 2020. Comparison of decay rates between native and non-native wood species in invaded forests of the southeastern U.S.: a rapid assessment.
• Ulyshen, Michael ; Horn, Scott ; Hanula, James L. 2020. Effects of Chinese privet on bees and their vertical distribution in riparian forests.
• Vogt, James T.; Coyle, David R.; Jenkins, David ; Barnes, Chris ; Crowe, Christopher ; Horn, Scott ; Bates, Chip ; Roesch, Francis A. 2020. Efficacy of five herbicide treatments for control of Pyrus calleryana.
• Poole, Emilee M.; Ulyshen, Michael D.; Horn, Scott ; Cram, Michelle ; Olatinwo, Rabiu ; Fraedrich, Stephen . 2019. Biology and distribution of Agrilus macer LeConte (Coleoptera: Buprestidae), a species associated with sugarberry (Celtis laevigata Willd.) mortality in the southeastern USA.
• Ulyshen, Michael ; Cram, Michelle ; Horn, Scott ; Villari, Caterina ; Poole, Emilee . 2019. Sugarberry dieback and mortality.
• Sheehan, Thomas N.; Ulyshen, Michael D.; Horn, Scott ; Hoebeke, E. Richard. 2019. Vertical and horizontal distribution of bark and woodboring beetles by feeding guild: is there an optimal trap location for detection?.
• Ulyshen, Michael D.; Horn, Scott ; Pokswinski, Scott ; McHugh, Joseph V; Hiers, J. Kevin. 2018. A comparison of coarse woody debris volume and variety between old-growth and secondary longleaf pine forests in the southeastern United States.
• Ulyshen, Michael; Pawson, Stephen; Branco, Manuela; Horn, Scott; Hoebeke, Richard; Gossner, Martin. 2018. Utilization of non-native wood by saproxylic insects.
• Ulyshen, Michael; Pawson, Stephen; Branco, Manuela; Horn, Scott; Hoebeke, Richard; Gossner, Martin. 2018. Utilization of non-native wood by saproxylic insects.
• Ulyshen, Michael D.; Shefferson, Richard; Horn, Scott; Taylor, Melanie K.; Bush, Bryana; Brownie, Cavell; Seibold, Sebastian; Strickland, Michael S. 2017. Below- and above-ground effects of deadwood and termites in plantation forests.
• Hanula, James L.; Ulyshen, Michael D.; Horn, Scott. 2016. Conserving pollinators in North American forests: A review.
• Zhang, Yanzhuo; Hanula, James L.; Horn, Scott; Jones, Cera; Braman, S. Kristine; Sun, Jianghua. 2016. Fundamental host range of Leptoypha hospita (Hemiptera: Tingidae), a potential biological control agent of Chinese privet.
• Hanula, James L.; Mayfield, Albert (Bud); Reid, Laurie S.; Horn, Scott. 2016. Influence of trap distance from a source population and multiple traps on captures and attack densities of the redbay ambrosia beetle (Coleoptera: Curculionidae: Scolytinae).
• Maner, M. Lake; Hanula, James; Horn, Scott. 2014. Population trends of the redbay ambrosia beetle (Coleoptera: Curculionidae: Scolytinae): does utilization of small diameter redbay trees allow populations to persist?.
• Zhang, Yanzhuo; Hanula, James; O'Brien, Joseph; Horn, Scott; Kris, Braman; Sun, Jianghua. 2013. Evaluation of the impacts of herbivory by lace bugs on Chinese privet (Ligustrum sinense) survival and physiology.
• Mayfield, Albert (Bud), III.; MacKenzie, Martin; Cannon, Philip G.; Oak, Steve; Horn, Scott; Hwang, Jaesoon; Kendra, Paul E. 2013. Suitability of California bay laurel and other species as hosts for the non-native redbay ambrosia beetle and granulate ambrosia beetle.
• Moorman, Christopher E.; Woen, Liessa T.; Kilgo, John C.; Hanula, James L.; Horn, Scott; Ulyshen, Michael D. 2012. Arthropod abundance and seasonal bird use of bottomland forest harvest gaps.
• Burke, Jordon L.; Hanula, James L.; Horn, Scott; Audley, Jackson P.; Gandhi, Kamal JK. 2012. Efficacy of two insecticides for protecting loblolly pines (Pinus taeda L.) from subcortical beetles (Coleoptera: Curculionidae and Cerambycidae).
• Ulyshen, Michael D.; Horn, Scott; Hanula, James L. 2012. Growth responses of mature loblolly pine to dead wood manipulations.
• Zhang, Yanzhuo; Hanula, James L.; Horn, Scott; Braman, Kristine; Sun, Jianghua. 2011. Biology of Leptoypha hospita (Hemiptera: Tingidae), a Potential Biological Control Agent of Chinese Privet.
• Hanula, James; Horn, Scott; Taylor, John W. 2010. Chinese Privet (Ligustrum sinense) removal and its effect on native plant communities of Riparian Forests.
• Ulyshen, Michael; Horn, Scott; Hanula, James. 2010. First Southeast U.S. records for Lacconotus punctatus Leconte (Coleoptera: Mycteridae), an early seasonal canopy specialist?.
• Hanula, James L.; Horn, Scott. 2009. Impact of Chinese privet and its removal on pollinator diversity and abundance.
• Horn, Scott; Ulyshen, Michael. 2009. The importance of streamside sandbars to ground beetle (Coleoptera, Carabidae) communities in a deciduous forest.
• Horn, S.; Ulyshen, M.D. 2009. The importance of streamside sandbars to ground beetle (Coleoptera, Carabidae) communities in a deciduous forest.
• Horn, Scott; Hanula, James L. 2008. Relationship of coarse woody debris to arthropod availability for red-cockaded woodpeckers and other bark-foraging birds on loblolly pine boles.
• Ulyshen, Michael D.; Hanula, James L.; Horn, Scott. 2007. Burying beetles (Coleoptera: Silphidae) in the forest canopy: The unusual case of Nicrophorus pustulatus Herschel.
• Klepac, John; Rummer, Robert B.; Hanula, James L.; Horn, Scott. 2007. Mechanical removal of Chinese privet.
• Moorman, Christopher E.; Bowen, Liessa T.; Kilgo, John C.; Sorenson, Clyde E.; Hanula, James L.; Horn, Scott; Ulyshen, Mike D. 2007. Seasonal diets of insectivorous birds using canopy gaps in a bottomland forest.
• Horn, Scott; Hanula, James L. 2006. Burlap bands as a sampling technique for green anoles (Anolis carolinensis) and other reptiles commonly found on tree boles.
• Horn, Scott; Hanula, James L. 2006. Burlap bands as a sampling technique for green anoles (Anolis carolinensis) and other reptiles commonly found in tree boles.
• Horn, Scott; Horn, George N. 2006. New host record for the Asian Ambrosia beetle, Xylosandrus crassiusculus (Motschulsky) (Coleoptera: Curculionidae).
• Hanula, James L.; Horn, Scott; Wade, Dale D. 2006. The role of dead wood in maintaining arthropod diversity on the forest floor.
• Horn, Scott; Hanula, James L.; Ulyshen, Michael D.; Kilgo, John C. 2005. Abundance of green tree frogs and insects in artificial canopy gaps in a bottomland hardwood forest.
• Ulyshen, Michael D.; Hanula, James L.; Horn, Scott; Kilgo, John C.; Moorman, Christopher E. 2005. Herbivorous insect response to group selection cutting in a southeastern bottomland hardwood forest.
• Ulyshen, Michael D.; Hanula, James L.; Horn, Scott; Kilgo, John C.; Moorman, Christopher E. 2005. The response of ground beetles (Coleoptera: Carabidae) to selection cutting in a South Carolina bottomland hardwood forest.
• Ulyshen, Michael D.; Hanula, James L.; Horn, Scott. 2005. Using Malaise traps to sample ground beetles (Coleoptera: Carabidae).
• Horn, Scott; Hanula, James L. 2004. A Survey of Cavity-Nesting Bees and Wasps in Loblolly Pine Stands of the Savannah River Site, Aiken County, South Carolina.
• Hanula, James L.; Horn, Scott. 2004. Availability and abundance of prey for the red-cockaded woodpecker.
• Horn, Scott; Hanula, James L. 2004. Impact of Seed Predators on the Herb Baptisia Lanceolata (Fabales: Fabaceae).
• Ulyshen, Michael D.; Hanula, James L.; Horn, Scott; Kilgo, John C.; Moorman, Christopher E. 2004. Spatial and temporal patterns of beetles associated with coarse woody debris in managed bottomland hardwood forests.
• Horn, Scott; Hanula, James L. 2002. Comparison of arthropod prey of red-cockaded woodpeckers on the boles of long-leaf and loblolly pines.
• Horn, Scott; Hanula, James L. 2002. Life History and Habitat Associations of the Broad Wood Cockroach, Purcoblatta lata (Blattaria: Blattellidae) and Other Native Cockroaches in the Coastal Plain of South Carolina.

Research Highlights

Assessing the Role of a Little-known Wood-boring Beetle in Sugarberry Decline (2019)

Ongoing research seeks to explain an ongoing, severe episode of sugarberry mortality in the southeastern U.S. A rarely collected species of buprestid beetle, Agrilus macer, is attacking dying trees at high densities. USDA Forest Service research results suggest that Agrilus macer is a secondary pest, rather than a primary cause of mortality, and there is no evidence that it is the vector for harmful pathogens.

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